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Answering questions over enterprise and scientific data lakes requires systems to navigate heterogeneous, weakly structured collections of tables, passages, and linked metadata. Current benchmarks abstract away this noisy discovery process, failing to evaluate end-to-end performance. To bridge this gap, we introduce LakeQuest, a human-validated benchmark of 9,846 QA pairs designed to evaluate the end-to-end retrieve-and-synthesize pipeline over realistic data lakes. LakeQuest spans three diverse domains (AI/ML metadata, retail banking, and multimodal biomedical drug information) and pairs every question with exact, modality-aware evidence pointers. By isolating source discovery from cross-modal synthesis, LakeQuest exposes critical failure modes in modern QA systems. 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